Albatrosses |
The true 'Ocean Wanderer'! A Northern
Royal Albatorss soars without effort on its huge wings.
| Systematics and evolution
of the Albatrosses
Until recently, the albatrosses have divided into two or three groups consisting of the so-called 'Great' albatrosses (Diomedea), the medium-sized Northern Pacific albatrosses (Phoebastria), the medium-sized Southern 'Mollymawks' (Thalassarche) and the exquisite Sooty Albatrosses (Phoebetria). Historically, there has been much confusion due to the frequent collection of albatrosses at sea. What were thought to be distinct forms (i.e. new species or subspecies) often turned out to be immature stages of known species. Recently, Gary Nunn and colleagues used mitochondrial cytochrome-b sequences to re-evaluate the phylogenetic relationships within the albatrosses (Nunn et al. 1996). This study concluded that the traditional genus Diomedea is in fact paraphyletic, thus supporting the four subgroups outlined above. Robertson and Nunn (1998) urge for a generous rather than conservative approach towards albatross systematics as a way to maximize conservation efforts. Obviously numerically limited 'taxa' will benefit in terms of conservation effort by being distinguised from more abundant taxa. Russ and Shirihai (2000) treat the suggested splits as allospecies rather than full species. Identification: Albatross identification is complicated by geographical, age-related and sex-related variation (see individual accounts for more information). Range: It is often said that albatrosses are restricted to the oceans of the southern hemisphere because of their need for wind. This is simply not true. Firstly, albatrosses are common in the north Pacific, where three species are endemic. Secondly, the fossil record indicates that albatrosses once enjoyed a more cosmopolitan distribution and extinct albatrosses are known from many northern hemisphere locations including North Carolina, England, Bermuda and California. Thirdly, southern albatrosses regularly wander into the North Atlantic and North Pacific. |
Diomedea exulans Snowy
[Wandering] Albatross (VULNERABLE)
Monotypic
Also known as Snowy Albatross
or White-winged Albatross
Diomedea dabbenena Tristan
Albatross (new candidate for consideration)
Monotypic
Formerly considered subspecies
of Wandering Albatross
Diomedea antipodensis Antipodean
Albatross (new candidate for consideration)
Monotypic
Formerly considered subspecies
of Wandering Albatross
Diomedea gibsoni Gibson's
Albatross (new candidate for consideration)
Monotypic
Formerly considered subspecies
of Wandering Albatross
Diomedea amsterdamensis Amsterdam
Albatross (CRITICAL)
Monotypic
A pair of Southern Royal
Albatross nesting amongst wind-swept tussock grass on Campbell Island,
New Zealand. Photo copyright of Angus Wilson.
Diomedea epomophora Southern
Royal Albatross (NEAR THREATENED)
Monotypic
Diomedea sanfordi Northern
Royal Albatross (new candidate for consideration)
Monotypic
Diomedea irrorata Waved
Albatross (NEAR THREATENED)
Monotypic
Also known as Galapagos
Albatross
Diomedea albatrus Short-tailed
Albatross (ENDANGERED)
Monotypic
Also known as Steller's
Albatross
Diomedea nigripes Black-footed
Albatross (new candidate for consideration)
Monotypic
Diomedea immutabilis Laysan
Albatross
Monotypic
Diomedea melanophrys Black-browed
Albatross (new candidate for consideration)
Now monotypic
Formerly genus Thalassarche
Also known as Black-browed
Mollymawk
Diomedea impavida Campbell
Albatross (new candidate for consideration)
Monotypic
Formerly subspecies of Black-browed
Albatross.
Formally genus Thalassarche
Diomedea cauta Shy
Albatross (new candidate for consideration)
Now monotypic. Also known
as Tasmanian Shy Albatross.
Formerly genus Thalassarche
Diomedea steadi White-capped
Albatross (new candidate for consideration)
Monotypic
Formerly subspecies of Shy
Albatross. Also known as Auckland Shy Albatross.
Formerly genus Thalassarche
Diomedea salvini Salvin's
Albatross (new candidate for consideration)
Monotypic
Formerly subspecies of Shy
Albatross.
Formerly genus Thalassarche
Also known as Bounty Albatross
or Grey-backed Albatross
Diomedea eremita Chatham
Albatross (new candidate for consideration)
Monotypic
Formerly subspecies of Shy
Albatross.
Formerly genus Thalassarche
Diomedea chrysostoma Grey-headed
Albatross (NEAR THREATENED)
Monotypic
Formerly genus Thalassarche
Also known as Flat-billed
Albatross, Gould's Albatross, Grey-mantled Albatross, Yellow-nosed Albatross(!),
and Grey-headed Mollymawk.
Diomedea chlororhynchos Atlantic
Yellow-nosed Albatross (new candidate for consideration)
Monotypic
Formerly genus Thalassarche
Also known as Carter's Albatross(?)
Diomedea carteri Indian
Yellow-nosed Albatross (new candidate for consideration)
Monotypic
Formerly genus Thalassarche
Diomedea bulleri Buller's
Albatross (new candidate for consideration)
Monotypic
Formerly genus Thalassarche
Also known as Buller's Mollymawk
Diomedea nov.sp.? Pacific
Albatross (new candidate for consideration)
Monotypic
Formerly the platei subspecies
of Buller's Albatross.
Formerly genus Thalassarche
Phoebetria fusca Sooty
Albatross (NEAR THREATENED)
Monotypic
Also known as Dark-mantled
Sooty Albatross
Phoebetria palpebrata Light-mantled
Albatross (new candidate for consideration)
Monotypic
Also known as Light-mantled
Sooty Albatross or Grey-mantled Albatross
Enticott, J. and Tipling, D. (1997) Photographic Guide to Seabirds of the World. New Holland, London.
Nunn, G. B. and Stanley, S. E. (1998) Body size effects and rates of cytochrome b evolution in tube-nosed seabirds. Mol. Biol. Evol., 15(10): 1360-1371.
Nunn, G. B., Cooper, J., Jouventin, P., Robertson, C. J. R. and Robertson G. G. (1996) Evolutionary relationships among extant albatrosses (Procellariiformes: Diomedeidae) established from complete cytochrome-b gene sequences. Auk 113: 784-801.
Robertson, C. J. R. and Nunn, G. B. (1998) Towards a new taxonomy for albatrosses. In 'Albatross Biology and Conservation'. (Eds. Robertson, G. and Gales, R.) Chp 2, p 13-19. Surrey Beatty and Sons Ltd.
Robertson, G. and Gales, R. (1998) Albatross Biology and Conservation. Surrey Beatty and Sons Ltd. (ISBN 0 949324 82 5).
Russ, R. and Shirihai, H. (2000) The birds, marine mammals, habitat and history of the subantarctic islands off New Zealand. Alula 3(6): 82-147.
Shirihai, H. and Jarrett, B. (2002) A complete Guide to Antarctic Wildlife: the birds and marine mammals of the Antarctic Continent and Southern Ocean. Alula Press, Degerby, Finland.
Tickell, W. L. N. (2000) Albatrosses. Yale University Press.
Tickell, W.L.N. (1970) The great albatrosses. Scientific American 223(5): p84-93.
Copyright © 2002 All
rights reserved. Angus Wilson
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